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3-hydroxykynurenine hydroxykynurenine metabolite tryptophan uv light xanthurenic acid kyn levels kmo total serum hydroxyanthranilic amino quinolinic l-3-hk 3hk bioprecursor endogenous excitotoxin radical generator 3d rendering chemical biochemistry molecule molecular structural structure formula skeletal isomer production metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic hmdb intermediate precursor synthesis derivative