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3-methylglutaconyl-coa mg-coa hydratase deficiency fatty acid amino acp tissue distribution isoprenoids methylcrotonyl coa carboxylase methylglutaconic aciduria type catabolism acinetobacter isovaleryl hmg leucine hmg-coa 3d rendering chemistry molecule molecular structural structure formula skeletal geometry metabolite metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor