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chlorophyll oxygenic photosynthesis violet-blue orange-red universal eukaryotes land plants prochlorophytes antenna complex photosynthetic organisms anoxygenic chlorophyl chlorophyll pigment cyanobacteria chloroplasts algae energy light electromagnetic spectrum wavelengths photosystems thylakoid resonance energy transfer reaction center absorb photosystem ii electron transport chain light nadph porphyrin chlorin magnesium 3d phytyl biosynthesis chlorophyllide photosensitizer cycle molecule molecular formula skeletal structural structure