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vitamin b6 pyridoxal 5-phosphate phosphate vitamin essential nutrient vitamer coenzyme amino acid glucose lipid ultraviolet-b radiation chlorophyll anti-vitamin neurotransmitter synthesis histamine hemoglobin synthesis gene expression macronutrient metabolism decarboxylation transamination racemization elimination replacement metabolite biosynthesis cycle mechanism catabolism biosynthetic intermediate biogenesis substrate precursor derivative bioprecursor metabolic enzyme endogenous contractor molecule molecular formula skeletal structural structure geometry 3d