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glucosamine 2-amino-2-deoxy-glucose amino sugar glycosylated proteins metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate precursor synthesis derivative metabolic endogenous polysaccharides chitosan chitin monosaccharides shellfish exoskeletons dietary supplement biological activity bioactive nutrient prescription drug food additive nutraceutica dietary arthritis pain chondroitin chondroitin sulfate methylsulfonylmethane blood thinner warfarin aspergillus niger fermenting corn molecule molecular formula skeletal structural structure 3d