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glyceraldehyde 3-phosphate triose phosphate 3-phosphoglyceraldehyde 2-hydroxy-3-oxopropyl dihydrogen g3p ga3p gadp gap tp galp pgal metabolite metabolism biosynthesis cycle pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor pentose phosphate gluconeogenic photosynthesis tryptophan thiamine molecule molecular formula skeletal structural structure geometry ball stic cell biology biochemistry biomolecule bioactive 3d