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glycerol 3-phosphate phosphate g3p phosphoric ester glycerol glycerophospholipid 3-o-phosphonoglycerol 3-phosphoglycerol gro3p l-glycerol d-glycerol 1-phosphate glycerate glyceraldehyde dhap glycolysis intermediate citric acid cycle glycolysis pathway mitochondrial membrane cytosol substrate enzyme phosphoglycolate phosphatase pgp g3pp enantiomer 3d metabolite metabolism biosynthesis mechanism path catabolism biosynthetic biogenesis precursor synthesis derivative bioprecursor metabolic endogenous contractor human metabolome database hmdb