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galactose gal monosaccharide sugar brain sweet epimer glucose lactose galactan hemicellulose nerve tissue pyranose furanose stereocenter tautomer isomer stereoisomer hydrolysis enzymes lactase dairy products avocados sugar beets glycolipids glycoproteins substance matter material compound component agent stuff reagent functional group precursor synthesis reaction production building block molecule molecular formula skeletal structural structure biochemistry biomolecule organic 3d