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uridine diphosphate glucose uracil-diphosphate udp-glucose nucleotide sugar glycosyltransferase sugar metabolism glycogen udp-galactose udp-glucuronic acid polysaccharides galactose glucuronic sucrose lipopolysaccharides glycosphingolipids pyrophosphate group ribose uracil metabolite biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb glucosyltransferases 3d molecule molecular formula skeletal structural structure