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galactose 1-phosphate intermediate d-galactose-1-phosphate glucose uridine diphosphate galactose galactokinase metabolism galactosemia leloir pathway enzyme genetic disorder galt galactose-1-phosphate uridylytransferase udp-activator group cataract deficiency test 3d metabolite biosynthesis cycle mechanism path catabolism biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor human metabolome database hmdb bioactive cell biochemistry biology biomolecule