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uridine diphosphate galactose metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb udp-galactose polysaccharide nucleotide sugars transferase b4galt5 molecule molecular formula skeletal structural structure geometry ball stick 3d isolated white background model bioactive cell biochemistry biomolecule biology substance matter compound