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chenodeoxycholic acid chenodesoxycholic chenocholic bile carboxylic chenodeoxycholate bile cheno cholesterol taurine glycine taurochenodeoxycholate glycochenodeoxycholate lipid steroid steroidal endogenous metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme contractor gallstones cerebrotendineous xanthomatosis hepatitis molecule molecular formula skeletal structural structure 3d