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taurocholic acid cholaic cholyltaurine acidum cholatauricum bile emulsification fats bile conjugate cholic taurine cholagogue choleretic cattle cholesterol precursor liver chenodeoxycholic ph metabolite metabolism biosynthesis cycle mechanism biosynthetic intermediate biogenesis substrate synthesis derivative bioprecursor metabolic enzyme endogenous biochemistry biomolecule bioactive biochemical molecule molecular formula skeletal structural structure 3d