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lithocholic acid lca bile detergent solubilize fats colon chenodeoxycholic steroid dietary fiber stool vitamin receptor anandamide lipid metabolite metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor phosphatidylcholine lecithin sphingomyelin phospholipid membrane choline lipid phosphocholine cholesterol liver bile steroid taurine glycine salts taurocholic glycocholic 3d