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metabolite diiodothyronine function action thyroid iodine deiodinase custom synthesis pharmaceutical diiodo thyronine hypothyroidism iodothyronine propanoic acid gland hyperthyroidism endocrine triiodothyronine level 3d rendering chemical biochemistry molecule molecular structural structure formula skeletal substance compound component production derivative hormone hormonal receptor agonist metabolism biosynthesis mechanism pathway path catabolism biosynthetic intermediate biogenesis