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serine 2-amino-3-hydroxypropanoic acid ser amino proteins group metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous proteinogenic stereoisomer tautomer isomer essential glycine silk protein signaling neurons neuromodulator nmda receptors alzheimer's disease biomarker cerebrospinal fluid biomolecule bioactive biochemical molecule molecular formula skeletal structural structure 3d