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mechanism phosphoenolpyruvate structure reaction oxaloacetate biology 2-phosphoglycerate phosphoglycerate 2pg 2-phosphoglyceric acid phosphoglyceric glycolysis serine glycogen glucose phosphate atp tca glycolytic pathway pep glyceric substrate step enolase partial breakdown lactate oxidation 3d rendering chemical biochemistry molecule molecular structural formula compound synthesis derivative metabolite metabolism biosynthesis cycle path catabolism biosynthetic intermediate