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2-aminomuconic semialdehyde glutamate carboxymuconate acid intermediate tryptophan ball stick degradation compound dehydrogenase aminophenol kynurenine muconic tryptophan-niacin catabolic zwitterion amino trp aminomuconic muconate 3d rendering chemistry chemical biochemistry biology atom molecule molecular structural structure formula skeletal geometry component reaction synthesis derivative metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic