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glycocyamine guanidinoacetate 2-guanidinoacetic acid metabolite glycine guanidine guanylation arginine creatine supplement feed additive poultry farming metabolism homocysteine serine threonine proline biochemical synthesis urea cycle gamt weight gain muscle increase betaine rendering biosynthesis mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb