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isopentenyl pyrophosphate ipp diphosphate idp hydroxy- 3-methylbut-3-enoxy phosphoryl oxyphosphonic acid isoprenoid precursor hmg-coa reductase pathway mevalonate non-mevalonate mep biosynthesis isomer dmapp terpenes terpenoids acetyl-coa dimethylallyl pyrophosphate isomerase hmb-pp hmb-pp reductase plastids bacteria apicomplexan protozoa rendering metabolite metabolism cycle mechanism path catabolism biosynthetic intermediate biogenesis substrate synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb