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dimethylallyl pyrophosphate dmapp diphosphate dmadp isoprenyl isoprenoid precursor mevalonate pathway mep biosynthesis ipp isomer isopentenyl pyrophosphate isomerase mevalonic acid terpenes terpenoid plant isoprenoid hmg-coa reductase eukaryotes archaea bacteria rendering metabolite metabolism cycle mechanism path catabolism biosynthetic intermediate biogenesis synthesis derivative bioprecursor metabolic enzyme endogenous contractor bioactive cell biochemistry biology biomolecule