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homovanillic acid hva catecholamine 4-hydroxy-3-methoxy-benzeneacetic 3-methoxy-4-hydroxyphenylacetic monoamine oxidase catechol-o-methyltransferase dopamine oxidative enzymes psychiatry cerebrospinal fluid metabolic stress 2-deoxy-d-glucose neurotransmitter receptor agonist neuromodulatory nervous system brain appetite memory signaling neuroscience neuroblastoma pheochromocytoma post-menopausal transgender people cross-sex hormone tobacco metabolite metabolism biosynthesis pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous 3d