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fucoxanthin xanthophyll pigment chloroplast brown algae heterokonts olive-green plant photosynthesis beta-carotene fucus spp microalgae unicellular diatoms macroalgal plastid photosystem chlorophyll seaweeds fucus dictyota laminaria photobioreactor phaeodactylum tricornutum dietary ingredient ndi 3d metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb