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diiodotyrosine dit precursor thyroid hormone iodization monoiodotyrosine phenol ring enzyme peroxidase triiodothyronine follicle rendering thyroid iodine deiodinase hypothyroidism gland hyperthyroidism thyromimetic thermogenesis hormonal therapy replacement function action endocrine level prohormone analogue metabolite metabolism biosynthesis cycle mechanism pathway catabolism biosynthetic intermediate biogenesis substrate synthesis derivative bioprecursor metabolic endogenous contractor