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ceramide waxy lipid sphingosine fatty acid phosphatidylcholine pc lecithin sphingomyelin phosphatidylethanolamine phospholipid phosphocholine lysophosphatidylcholine surfactant glycerophospholipid phosphatidylserine lipophilic cell membrane lipid bilayer sphingolipid cellular signaling differentiation proliferation programmed death pcd vernix caseosa synthesis hydrolysis salvage pathway apoptosis skin epidermis hormonal leptin insulin resistance metabolite metabolism biosynthesis cycle mechanism biosynthetic intermediate biogenesis substrate bioprecursor endogenous 3d