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dipalmitoylphosphatidylcholine dppc phospholipid phosphatidylcholine pc lecithin sphingomyelin phosphatidylethanolamine membrane choline cell arachidonic acid lipid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic pulmonary alveolar collapse breathing liposome lung ls molecule molecular formula skeletal structural structure geometry ball stick biology biochemistry biomolecule bioactive metabolite metabolism biosynthesis cycle 3d