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ganglioside glycosphingolipid ceramide oligosaccharide sialic acid n-acetylneuraminic nana sugar chain globoside ganglion cells cell plasma membrane signal transduction lipid rafts immunology semisynthetic neurodegenerative phospholipids pituitary glycoprotein hormones metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor molecule molecular formula skeletal structural structure geometry biochemistry biomolecule bioactive 3d