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ceramide phosphatidylcholine pc lecithin sphingomyelin phosphatidylethanolamine phospholipid membrane choline cell arachidonic acid lipid phosphocholine cholesterol fatty lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic cell-permeable neurite mitochondrial respiratory chain waxy lipid cell cellular signaling differentiation proliferation programmed death 3d metabolite metabolism biosynthesis mechanism biosynthetic intermediate biogenesis substrate precursor derivative bioprecursor metabolic enzyme endogenous