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cardiolipin calcutta antigen component mitochondrial membrane lipid bacteria cl enzymes energy metabolism diphosphatidylglycerol phosphatidic acid moieties glycerol oxidative phosphorylation eukaryotic prokaryotic pathway catabolism anti-cardiolipin rendering phosphatidylserine precursor biosynthesis cycle surfactant acid cell phosphocholine choline phosphatidylethanolamine contractor enzyme hmdb lipophilic substrate cholesterol biosynthetic bioprecursor pc biogenesis lecithin bilayer pathway