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phosphatidylglycerol pulmonary surfactant lipid-gated ion channels alveolar phospholipase cardiolipin synthase cardiolipin mitochondrial membrane mitochondria cellular surface phosphomonoester phosphatidylcholine lecithin sphingomyelin phosphatidylethanolamine phospholipid cell arachidonic acid lipid phosphocholine fatty lysophosphatidylcholine bilayer glycerophospholipid phosphatidylserine lipophilic metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous 3d