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bisphosphatidylglycerol glycerophospholipid pulmonary surfactant plasma membrane cardiolipin mitochondrial 3d arachidonic acid mechanism endogenous sphingomyelin phosphocholine derivative lipophilic catabolism biosynthesis lipid hmdb ceramide phosphatidylethanolamine substrate phosphatidylcholine phosphatidylserine biosynthetic metabolism intermediate choline contractor enzyme phospholipid lecithin pathway bilayer cycle bioprecursor pc human metabolome database fatty path cholesterol biogenesis lysophosphatidylcholine synthesis