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citicoline cytidine diphosphate choline cdp-choline phosphatidylcholine cell membrane cells dietary supplement intestine ctp-phosphocholine cytidylyltransferase cognitive memory cognition ischemic stroke vision neuroprotective cardiolipin sphingomyelin arachidonic acid phosphatidylethanolamine neuronal signalling glutamate transport acetylcholine norepinephrine dopamine neurotransmitter rendering precursor contractor substrate lysophosphatidylcholine biosynthesis phosphatidylserine synthesis metabolic intermediate catabolism cycle biosynthetic mechanism derivative enzyme ceramide path arachidonic sphingomyelin