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biliverdin green tetrapyrrolic bile pigment heme catabolism bruises metabolite metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor heme hemoglobin erythrocytes hemosiderin bilirubin fluorescent cyanobacterial phycobiliprotein protein disease colour reductase function urine urine analysis molecule molecular formula skeletal structural structure 3d