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mesobilirubin bilirubin alpha heme catabolism pigment bile acid bilirubin catabolic red blood cells hemoglobin porphyrin heme biliverdin stercobilin urobilin urine tetrapyrrole pigment phycobilin phytochrome phototherapy glucuronyltransferase delta enterohepatic circulation hepatobiliary disease hyperbilirubinemia toxicity blood–brain barrier encephalopathy test levels metabolite metabolism biosynthesis mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous 3d