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bilirubin diglucuronide metabolism bilirubin hydrophilic hepatic br catabolic pathway heme vertebrates red blood cells hemoglobin rbcs erythrocytes porphyrin biliverdin tetrapyrrole glucuronic acid pigment phycobilin phytochrom isomer phototherapy newborn total alpha urine formation color rendering biosynthesis intermediate human metabolome database enzyme precursor substrate biogenesis path synthesis endogenous derivative catabolism biosynthetic cycle contractor metabolic