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a-aminoadipic acid alpha-aminoadipic conjugate a-aminoadipate a-aminoadipate pathway biochemical amino l-lysine euglenid fungi thermus metabolite metabolism biosynthesis cycle mechanism path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor homoisocitrate homoisocitrate dehydrogenase molecule molecular formula skeletal structural structure geometry ball stick bioactive cell biochemistry biomolecule biology chemistry chemical 3d