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n-acetylaspartylglutamic acid n-acetylaspartylglutamate naag third-most-prevalent neurotransmitter naa n-acetylaspartic spaglumic isospaglumic n-acetyl-1-aspartylglutamic n-acetyl-l-a-aspartyl-l-glutamic neurons naag peptidases synapse antiallergic biosynthesis neuroblastoma protein kinase catabolism amyotrophic lateral sclerosis hippocampus schizophrenia phencyclidine metabolite metabolism cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor molecule molecular formula skeletal 3d