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n-acetylneuraminic acid nana neu5ac sialic n-glycolylneuraminic cell membrane glycans mucins glycoproteins gangliosides neuronal membranes brain mucus mucous influenza hemagglutinin metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor molecule molecular formula skeletal structural structure geometry ball stick bioactive cell biochemistry biomolecule 3d