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6-phosphogluconolactone phosphogluconolactone pentose phosphate pathway phosphogluconate hexose monophosphate shunt nadph glucose ppp fischer reaction ribulose structure dehydration gluconic acid symbol glucosyl triphosphate deficiency fate isomerase 3d rendering chemical biochemistry molecule molecular structural formula metabolite metabolism biosynthesis cycle mechanism path catabolism biosynthetic metabolome hmdb intermediate substrate precursor synthesis metabolic enzyme