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uzarigenin coronilla valentina asclepias tuberosa cardenolide aglycon digitoxin cerberigenin echujetin evonogenin thevetigenin urarigenin odorigeni herbal phytochemical phytotherapy plant herb natural herbalism pharmacognosy pharmacopoeia molecule molecular formula skeletal structural structure geometry ball stick atom cell biology biochemistry biomolecule bioactive biochemical 3d metabolite biosynthesis mechanism biosynthetic intermediate biogenesis substrate synthesis derivative bioprecursor metabolic endogenous