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phycourobilin plant pigment tetrapyrrole orange photosynthesis cyanobacteria red algae phycobiliprotein phycoerythrin light-harvesting system absorption chromophore phycoerythrobilin phycobilisome synechococcus herbal phytochemical phytotherapy plant herb natural herbalism pharmacognosy pharmacopoeia metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate bioprecursor metabolic enzyme endogenous molecule molecular formula skeletal structural structure 3d