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phosphoribosylamine pra biochemical intermediate purine nucleotide inosine-5-monophosphate dna rna vitamin thiamine cobalamin enzyme amidophosphoribosyltransferase glutamine phosphoribosyl pyrophosphate prpp 3d metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor human metabolome database hmdb molecule molecular formula skeletal structural structure ball stick bioactive cell biochemistry