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glycineamide ribonucleotide gar biochemical intermediate purine nucleotides inosine-5-monophosphate dna rna thiamine cobalamin phosphoribosylamine pra formylglycinamide ribotide fgar phosphoribosylglycinamide formyltransferase 10-formyltetrahydrofolate rendering metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb bioactive cell biochemistry biology biomolecule molecule structural structure formula ball stick