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phosphoribosyl-n-formylglycineamide formylglycinamideribotide fgar formylglycinamide ribonucleotide ribotide biochemical intermediate purine nucleotides inosine-5-monophosphate dna rna vitamin thiamine cobalamin enzyme 3d metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor human metabolome database hmdb molecule molecular formula skeletal structural structure geometry ball stick atom