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popc phosphatidylcholine diacylglycerol phospholipid 1-palmitoyl-2-oleoylphosphatidylcholine palmitoyloleoylphosphatidylcholine lipid rafts eukaryotic cell membranes biophysical experiments rendering pc lecithin sphingomyelin phosphatidylethanolamine membrane choline cell arachidonic acid lipid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic metabolite metabolism biosynthesis mechanism pathway biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous