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lysophosphatidic acid lpa phospholipid derivative signaling molecule mitogen g-protein-coupled receptors cell proliferation autotaxin lpa gtpase activation stress fibers myosin light-chain phosphatase metabolism lysophospholipase phosphatidic 3d metabolite biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis precursor synthesis bioprecursor metabolic enzyme endogenous contractor phosphatidylcholine lecithin sphingomyelin phosphatidylethanolamine choline arachidonic phosphocholine cholesterol fatty ceramide lysophosphatidylcholine glycerophospholipid phosphatidylserine