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ketoglutaric acid 2-oxoglutaric 2-ketoglutaric alpha-ketoglutaric oxoglutaric glutaric krebs cycle metabolite metabolism biosynthesis mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor alanine transaminase amino acids nitrogen transporter brain detoxification reye's syndrome cirrhosis urea cycle disorder antioxidant immune regulation cell biology biochemistry biomolecule bioactive molecule molecular formula skeletal structural structure 3d