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insulin peptide hormone beta cells pancreatic islets ins gene islets langerhans pancreas endocrine metabolism carbohydrates glucose glycogen triglycerides lipogenesis secretion catabolism reserve body fat blood sugar levels insulin glycaemia concentration glucose level glucagon glycogenolysis gluconeogenesis homeostasis diabetes diabetes mellitus carbohydrate synthesis signal peptidase molecular formula human peptide transporter glycolytic pathway calcium ion channels glucose-dependent insulinotropic gip tolerance test 3d endocrinology metabolite biosynthesis mechanism