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estrin oestrin estratriene estrane steroid double bonds parent structure estrogen hormones estradiol estrone estriol derivative binding affinity estrogen receptor progesterone expression ovaries estrus rendering contraception birth control hormonal oral contraceptives mother gynecology obstetrics woman motherhood birth fertility reproduction reproductive female sex hormone menopausal hormone therapy gynecological conditions pregnancy hormonal replacement function action endocrine endocrinology level prohormone