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cystathionine intermediate cysteine synthesis transsulfuration pathway homocysteine cystathionine gamma-lyase cth cysteine dioxygenase cdo sulfinoalanine decarboxylase hypotaurine taurine glutamate–cysteine ligase gcl glutathione synthetase gss glutathione cystathioninuria beta synthase ketobutyrate 3d metabolite metabolism biosynthesis cycle mechanism path catabolism biosynthetic biogenesis substrate precursor derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb bioactive cell biochemistry biomolecule